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Library of Congress Cataloging‐in‐Publication Data
Names: Binkley, Dan, author. | John Wiley & Sons, publisher.
Title: Forest ecology : an evidence‐based approach / Dan Binkley, School of Forestry, Northern Arizona University.
Description: First edition. | Hoboken, NJ : Wiley‐Blackwell, 2021. | Includes index.
Identifiers: LCCN 2021001373 (print) | LCCN 2021001374 (ebook) | ISBN 9781119703204 (paperback) | ISBN 9781119704409 (adobe pdf) | ISBN 9781119704416 (epub)
Subjects: LCSH: Forest ecology.
Classification: LCC QH541.5.F6 B555 2021 (print) | LCC QH541.5.F6 (ebook) | DDC 577.3–dc23
LC record available at https://lccn.loc.gov/2021001373
LC ebook record available at https://lccn.loc.gov/2021001374
Cover Design: Wiley
Cover Image: © Dan Binkley
The development of forests always includes contingent events: if an event happens, such as a fire, windstorm, or insect outbreak, the future of the forest will unfold differently than if the event did not happen (or if it happened in some other way at another time). This book would not be in front of you without the contingent event of Wally showing up as a young professor when I was an undergraduate at the School of Forestry at Northern Arizona University. Wally's engaging curiosity, interest in students, and active research program pulled my interests and future path into the domain of forest ecology. He continued to be a mentor through my grad student days at other universities, and most recently he led us through establishing the Colorado Forest Restoration Institute (modeled on NAU's Ecological Restoration Institute). It's been a good path.Thanks Wally.
Dan Binkley Fort Collins, Colorado
Preface
How Do We Come to Understand Forests?
This book supports learning about forest ecology. A good place to start is with a few points about knowledge, followed by a framework on how to approach forest ecology, some key features of using graphs to interpret information, and finally coming around to how to think about questions and answers in forests.
Humans try to understand complex worlds through a range of perspectives. Art tries to capture some essential features of a complex world, emphasizing how parts interact to form wholes. Religions explain how worlds work now, how the worlds came to be, and what will come next. Both art and religion develop from ideas and concepts, originated by individual artists or passed down by religious societies. How do we know if a work of art or an idea in religion represents the real world accurately? This question generally isn’t important. Art that satisfies the artist is good art, and religions are accepted on faith.
Art and religion have been evolving for more than 100 000 years, and lands and forests have been part of that development. One of the first written stories is a religious one from the Epic of Gilgamesh, from more than 4000 years ago from the Mesopotamian city of Uruk (now within Iraq). Gilgamesh and a companion traveled to the distant, sacred Cedar Mountain to cut trees. Lines from the epic poem include (based on Al‐Rawi and George 2014):
They stood there marveling at the forest, observing the height of the cedars … They were gazing at the Cedar Mountain, dwelling of gods, sweet was its shade, full of delight. All tangled was the thorny undergrowth, the forest a thick canopy, cedars so entangled it had no ways in. For one league on all sides cedars sent forth saplings, cypresses for two‐thirds of a league. Through all the forest a bird began to sing … answering one another, a constant din was the noise. A solitary tree‐cricket set off a noisy chorus. A wood pigeon was moaning, a turtle dove calling in answer. At the call of the stork, the forest exults. At the cry of the francolin bird, the forest exults in plenty. Monkey mothers sing aloud, a youngster monkey shrieks like a band of musicians and drummers, daily they bash out a rhythm …
And after slaying the demigod who protected the forest, Gilgamesh's companion laments:
My friend, we have cut down a lofty cedar, whose top abutted the heavens … We have reduced the forest to a wasteland.
What would actually happen if cedar trees were cut on a mountain? Would more cedar trees establish, would the post‐cutting landscape provide suitable habitat for the birds and monkeys? Would floods result? Anything could happen next in a story, but understanding which stories about the real world warrant confidence depends on the strength of evidence.
The core of understanding is knowing how one thing connects to another, and if the connections are the same everywhere and all the time, or if local details strongly influence the connections. The seasonal movements of the sun across the sky are consistent across years, but appear to differ from southern to northern locations. Multiple stories might explain the Sun's march with reasonable accuracy. Patterns etched on rocks by ancient artists may line up with key points in the Sun's seasonal patterns, and the movements of the Sun may reliably follow ceremonies convened by a society with the goal of ensuring the Sun's path. With art and religion, people may have understood