And then when we pass to the compounds of organic chemistry, we find this general law still further exemplified: we find much greater complexity and much less stability. An atom of albumen, for instance, consists of 482 ultimate atoms of five different kinds. Fibrine, still more intricate in constitution, contains in each atom, 298 atoms of carbon, 40 of nitrogen, 2 of sulphur, 228 of hydrogen, and 92 of oxygen – in all, 660 atoms; or, more strictly speaking – equivalents. And these two substances are so unstable as to decompose at quite ordinary temperatures; as that to which the outside of a joint of roast meat is exposed. Thus it is manifest that the present chemical heterogeneity of the Earth's surface has arisen by degrees, as the decrease of heat has permitted; and that it has shown itself in three forms – first, in the multiplication of chemical compounds; second, in the greater number of different elements contained in the more modern of these compounds: and third, in the higher and more varied multiples in which these more numerous elements combine.
To say that this advance in chemical heterogeneity is due to the one cause, diminution of the Earth's temperature, would be to say too much; for it is clear that aqueous and atmospheric agencies have been concerned; and, further, that the affinities of the elements themselves are implied. The cause has all along been a composite one: the cooling of the Earth having been simply the most general of the concurrent causes, or assemblage of conditions. And here, indeed, it may be remarked that in the several classes of facts already dealt with (excepting, perhaps, the first), and still more in those with which we shall presently deal, the causes are more or less compound; as indeed are nearly all causes with which we are acquainted. Scarcely any change can with logical accuracy be wholly ascribed to one agency, to the neglect of the permanent or temporary conditions under which only this agency produces the change. But as it does not materially affect our argument, we prefer, for simplicity's sake, to use throughout the popular mode of expression.
Perhaps it will be further objected, that to assign loss of heat as the cause of any changes, is to attribute these changes not to a force, but to the absence of a force. And this is true. Strictly speaking, the changes should be attributed to those forces which come into action when the antagonist force is withdrawn. But though there is an inaccuracy in saying that the freezing of water is due to the loss of its heat, no practical error arises from it; nor will a parallel laxity of expression vitiate our statements respecting the multiplication of effects. Indeed, the objection serves but to draw attention to the fact, that not only does the exertion of a force produce more than one change, but the withdrawal of a force produces more than one change. And this suggests that perhaps the most correct statement of our general principle would be its most abstract statement – every change is followed by more than one other change.
Returning to the thread of our exposition, we have next to trace out, in organic progress, this same all-pervading principle. And here, where the evolution of the homogeneous into the heterogeneous was first observed, the production of many changes by one cause is least easy to demonstrate. The development of a seed into a plant, or an ovum into an animal, is so gradual, while the forces which determine it are so involved, and at the same time so unobtrusive, that it is difficult to detect the multiplication of effects which is elsewhere so obvious. Nevertheless, guided by indirect evidence, we may pretty safely reach the conclusion that here too the law holds.
Observe, first, how numerous are the effects which any marked change works upon an adult organism – a human being, for instance. An alarming sound or sight, besides the impressions on the organs of sense and the nerves, may produce a start, a scream, a distortion of the face, a trembling consequent upon a general muscular relaxation, a burst of perspiration, an excited action of the heart, a rush of blood to the brain, followed possibly by arrest of the heart's action and by syncope: and if the system be feeble, an indisposition with its long train of complicated symptoms may set in. Similarly in cases of disease. A minute portion of the small-pox virus introduced into the system, will, in a severe case, cause, during the first stage, rigors, heat of skin, accelerated pulse, furred tongue, loss of appetite, thirst, epigastric uneasiness, vomiting, headache, pains in the back and limbs, muscular weakness, convulsions, delirium, &c.; in the second stage, cutaneous eruption, itching, tingling, sore throat, swelled fauces, salivation, cough, hoarseness, dyspnœa, &c.; and in the third stage, œdematous inflammations, pneumonia, pleurisy, diarrhœa, inflammation of the brain, ophthalmia, erysipelas, &c.: each of which enumerated symptoms is itself more or less complex. Medicines, special foods, better air, might in like manner be instanced as producing multiplied results.
Now it needs only to consider that the many changes thus wrought by one force upon an adult organism, will be in part paralleled in an embryo organism, to understand how here also, the evolution of the homogeneous into the heterogeneous may be due to the production of many effects by one cause. The external heat and other agencies which determine the first complications of the germ, may, by acting upon these, superinduce further complications; upon these still higher and more numerous ones; and so on continually: each organ as it is developed serving, by its actions and reactions upon the rest, to initiate new complexities. The first pulsations of the fœtal heart must simultaneously aid the unfolding of every part. The growth of each tissue, by taking from the blood special proportions of elements, must modify the constitution of the blood; and so must modify the nutrition of all the other tissues. The heart's action, implying as it does a certain waste, necessitates an addition to the blood of effete matters, which must influence the rest of the system, and perhaps, as some think, cause the formation of excretory organs. The nervous connections established among the viscera must further multiply their mutual influences: and so continually.
Still stronger becomes the probability of this view when we call to mind the fact, that the same germ may be evolved into different forms according to circumstances. Thus, during its earlier stages, every embryo is sexless – becomes either male or female as the balance of forces acting upon it determines. Again, it is a well-established fact that the larva of a working-bee will develop into a queen-bee, if, before it is too late, its food be changed to that on which the larvæ of queen-bees are fed. Even more remarkable is the case of certain entozoa. The ovum of a tape-worm, getting into its natural habitat, the intestine, unfolds into the well-known form of its parent; but if carried, as it frequently is, into other parts of the system, it becomes a sac-like creature, called by naturalists the Echinococcus– a creature so extremely different from the tape-worm in aspect and structure, that only after careful investigations has it been proved to have the same origin. All which instances imply that each advance in embryonic complication results from the action of incident forces upon the complication previously existing.
Indeed, we may find à priori reason to think that the evolution proceeds after this manner. For since it is now known that no germ, animal or vegetable, contains the slightest rudiment, trace, or indication of the future organism – now that the microscope has shown us that the first process set up in every fertilized germ, is a process of repeated spontaneous fissions ending in the production