As, however, a general expression can be given to the variations referred to, and as they have an important geographical relationship, besides a general diagnosis, we give their characters and distribution in detail.
The general impression we derive from a study of the series is that the amount of white on the wing and elsewhere decreases from the Missouri River to the Pacific, exhibiting its minimum in Oregon and Washington, precisely as in the small black Woodpeckers; that in the Great Basin the size of the claws and the length of tail increases considerably; that the northern forms are entirely black, and the more southern brown or olivaceous, except on the head.
The following synopsis will be found to express the principal characteristics of the species and their varieties, premising that P. arcticus is more distinctly definable than any of the others. We add the character of the green-bodied Mexican species to complete the series.
1. Wing, 3.65; tail, 4.20. Outer tail-feather with terminal half of inner web white. Iris bright red, sometimes paler. Hab. Eastern Province United States. (Florida in winter.) … var. erythrophthalmus.
2. Wing, 2.90; tail, 3.75. Outer tail-feather with only terminal fourth of inner web white. Iris white. Hab. Florida (resident) … var. alleni.
A. Interscapulars with white streaks.
a. Outer webs of primaries not edged with white at the base.
1. Above olive-brown, the head and neck, only, continuous black; back streaked with black. White spots on wing-coverts not bordered externally with black. Wing, 3.25; tail, 4.00; hind claw, .44. Hab. Table-lands of Mexico … var. maculatus.21
2. Above black, tinged with olive on rump, and sometimes on the nape. White spots as in last. Inner web of lateral tail-feathers with terminal white spot more than one inch long; outer web broadly edged with white. Wing, 3.45; tail, 4.10; hind claw, .55. Female less deep black than male, with a general slaty-olive cast. Hab. Middle Province of United States, from Fort Tejon, California, to Upper Rio Grande, and from Fort Crook to Fort Bridger … var. megalonyx.
3. Above almost wholly black, with scarcely any olive tinge, and this only on rump. White spots restricted, and with a distinct black external border. White terminal spot on inner web of lateral tail-feather less than one inch long; outer web almost wholly black. Wing, 3.40; tail, 3.90; hind claw, .39. Female deep umber-brown, instead of black. Hab. Pacific Province of United States, south to San Francisco; West Humboldt Mountains … var. oregonus.
b. Outer webs of primaries distinctly edged with white at base.
4. Above black, except on rump, which is tinged with olivaceous. White spots very large, without black border. Inner web of lateral tail-feather with terminal half white, the outer web almost wholly white. Wing, 3.50; tail, 3.90; hind claw, .39. Female umber-brown, replacing black. Hab. Plains between Rocky Mountains and the Missouri; Saskatchewan Basin … var. arcticus.
B. Interscapulars without white streaks.
5. Above dusky olive; white spots on scapulars and wing-coverts small, and without black edge. Tail-patches very restricted (outer only .40 long). No white on primaries. Wing, 2.85; tail, 3.10. Female scarcely different. Hab. Socorro Island, off west coast of Mexico … var. carmani.22
PLATE XXXI.
1. Chondestes grammaca. ♂ Cal., 6300.
2. Pipilo erythrophthalmus. ♂ Pa., 2135.
3. Pipilo erythrophthalmus. ♀ Kansas, 8194.
4. Pipilo chlorura. ♂ Rocky Mts., 2644.
5. Pipilo arcticus. ♂ Dakota, 1944.
6. Pipilo arcticus. ♀.
7. Pipilo aberti. ♂ Ariz., 6748.
8. Pipilo crissalis. ♂ Cal., 5559.
9. Pipilo megalonyx. ♀.
10. Pipilo mesoleucus. ♂ Ariz., 6829.
11. Pipilo albigula. ♂ Cape St. Lucas, 12993.
12. Pipilo oregonus. ♀.
Fringilla erythrophthalma, Linn. Syst. Nat. I, 1766, 318.—Aud. Orn. Biog. I, 1832, 151; V, 511, pl. xxix. Emberiza erythrophthalma, Gm. Syst. Nat. I, 1788, 874.—Wilson, Am. Orn. VI, 1812, 90, pl. liii. Pipilo erythrophthalmus, Vieill. Gal. Ois. I, 1824, 109, pl. lxxx.—Bon. List, 1838.—Ib. Conspectus, 1850, 487.—Aud. Syn. 1839, 124.—Ib. Birds Am. III, 1841, 167, pl. cxcv.—Baird, Birds N. Am. 1858, 512.—Samuels, 333. Pipilo ater, Vieill. Nouv. Dict. XXXIV, 1819, 292. Towhee Bird, Catesby, Car. I, 34. Towhee Bunting, Latham, Syn. II, I, 1783, 199.—Pennant, II, 1785, 359.
2135 ♂
Sp. Char. Upper parts generally, head and neck all round, and upper part of the breast, glossy black, abruptly defined against the pure white which extends to the anus, but is bounded on the sides and under the wings by light chestnut, which is sometimes streaked externally with black. Feathers of throat white in the middle. Under coverts similar to sides, but paler. Edges of outer six primaries with white at the base and on the middle of the outer web; inner two tertiaries also edged externally with white. Tail-feathers black; outer web of the first, with the ends of the first to the third, white, decreasing from the exterior one. Outermost quill usually shorter than ninth, or even than secondaries; fourth quill longest, fifth scarcely shorter. Iris red; said to be sometimes paler, or even white, in winter. Length, 8.75; wing, 3.75; tail, 4.10. Bill black, legs flesh-color. Female with the black replaced by a rather rufous brown.
Hab. Eastern United States to the Missouri River; Florida (in winter).
The tail-feathers are only moderately graduated on the sides; the outer about .40 of an inch shorter than the middle. The outer tail-feather has the terminal half white, the outline transverse; the white of the second is about half as long as that of the first;