Passing from the evidence that Evolution has taken place to the question–How has it taken place?–Mr. Spencer finds in known agencies and known processes adequate causes of its phenomena. In astronomic, geologic, and meteorologic changes, ever in progress, ever combining in new and more involved ways, we have a set of inorganic factors to which all organisms are exposed; and in the varying and complicated actions of organisms on one another we have a set of organic factors that alter with increasing rapidity. Thus, speaking generally, all members of the Earth's flora and fauna experience perpetual rearrangements of external forces. Each organic aggregate, whether considered individually or as a continuously existing species, is modified afresh by each fresh distribution of external forces. To its pre-existing differentiations new differentiations are added; and thus that lapse to a more heterogeneous state, which would have a fixed limit were the circumstances fixed, has its limits perpetually removed by the perpetual change of the circumstances. These modifications upon modifications, which result in evolution, structurally considered, are the accompaniments of those functional alterations continually required to re-equilibrate inner with outer actions. That moving equilibrium of inner actions corresponding with outer actions, which constitutes the life of an organism, must either be overthrown by a change in the outer actions or must undergo perturbations that cannot end until there is a readjusted balance of functions and correlative adaptation of structures. But where the external changes are either such as are fatal when experienced by the individuals, or such as act on the individuals in ways that do not affect the equilibrium of their functions, then the readjustment results through the effects produced on the species as a whole: there is indirect equilibration. By the preservation in successive generations of those whose moving equilibria are less at variance with the requirements, there is produced a changed equilibrium completely in harmony with the requirements.
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