Bovine Reproduction. Группа авторов. Читать онлайн. Newlib. NEWLIB.NET

Автор: Группа авторов
Издательство: John Wiley & Sons Limited
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Жанр произведения: Биология
Год издания: 0
isbn: 9781119602460
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maintain a closed hydraulic system to contain the extraordinary pressures generated as the ischiocavernosus muscles rhythmically contract against the blood‐filled crura of the penis [2, 4]. If the integrity of the fibrous tunica albuginea is compromised, anastomoses between the CCP and the surrounding peripenile vasculature may form and provide an escape route for blood contained in the CCP. Should this occur, pressure sufficient to achieve or maintain erection cannot develop [28, 37]. Communication of the CCP with the corpus spongiosum penis will produce a similar result because venous drainage of the CSP is not occluded during erection [29, 38]. Formation of vascular shunts may follow traumatic disruption of the tunica albuginea or be associated with a congenital weakness of the tissues of the tunica albuginea.

Photos depict bull prepared for cavernosography.

      Source: Courtesy of Dwight Wolfe.

      Cases of erection failure due to multiple vascular shunts involving defects in the tunica albuginea of the free portion of the penis are thought to be the result of a congenital weakness in the structural integrity of the tunica. In such cases there is no history of penile trauma or injury. Affected bulls fail to achieve erection following sexual stimulation and may have bluish discoloration of the penile or preputial skin when attempting to breed or when attempts are made to induce erection with an electroejaculator. Discoloration is the result of blood exiting the CCP through the peripenile vasculature. Multiple distal vascular shunts are most often diagnosed in bulls during the first breeding season. Vascular shunts in the distal penis are readily demonstrable with cavernosography (see Figure 15.13). Unlike vascular shunts that form secondary to traumatic disruption of the tunica albuginea, surgical correction of multiple congenital shunts is unsuccessful.

      Filling Defects of the CCP

Photo depicts failure of the distal portion of the penis to become erect due to occlusion of the cavernous spaces of the corpus cavernosum penis.

      When mounting an estrus female the bull must position himself to make searching motions and use the penis to locate the vulva in order to make intromission. Sensory innervation to the glans and free portion of the penis is necessary for the bull to align the penis, achieve intromission, and successfully complete the breeding act. Without sufficient sensory innervation of the distal penis the bull is unable to locate the vagina and the coital act cannot be completed [41].

      Sensory input from the distal penis is transmitted by branches of the paired dorsal nerves of the penis through the pudendal nerves to reach the spinal cord and brain. Although disruption of any portion of the neurologic pathway could result in loss of sensation, damage to the dorsal nerves of the penis is the most likely etiology of penile desensitization. Denervation injury does not interfere with erection, but affected bulls are unable to breed by natural service and usually cannot be successfully collected with an AV. Semen collection with an electroejaculator for artificial insemination is possible.