Collins New Naturalist Library. L. Matthews Harrison

they include the wolf, the brown and cave bears Ursus arctos and U. spelaeus, a sabre-tooth Homotherium, the horse, giant deer, elk, a bison Bison priscus, and the aurochs.

      Man returned after the peak of the Devensian glaciation as shown by his artifacts and by a few skeletal remains.³⁰ Some human teeth of middle Devensian age from Picken’s Hole cave in Somerset are the earliest human remains known in the British Isles apart from the Swanscombe skull whose alleged age has been challenged by some people. The largest find of palaeolithic man belongs to the late Devensian deposits of Aveline’s Hole in the Mendip Hills of Somerset, where bones representing thirty-one skeletons were excavated.

      As the ice melted during the late glacial stage of the Devensian the climate became milder and reached a peak after about a thousand years in the Allerød interstadial or ‘amelioration’ as it is sometimes called, though amelioration could have a different meaning for a reindeer than for a red deer. Thereafter the climate again became colder until about 10,000 years ago when the ice finally disappeared inaugurating the Flandrian interglacial which has lasted until the present. By the end of the late glacial many of the large mammals had become extinct, although there appears to be no reason why they should not have survived into the Flandrian. Perhaps the change of climate and the resulting changes in vegetation deprived them of their ecological niches, but it is also possible that improved hunting skills of upper palaeolithic man may have overcropped and thus exterminated them. The few species not part of our present mammalian fauna that survived from the late glacial disappeared in the early part of the Flandrian, which is discussed in the next chapter.

       CHAPTER 3

       THE EVOLUTION OF THE ENVIRONMENT

      AT the beginning of the Flandrian stage, when the glaciation started to recede, the climate became warmer and thereafter varied between warmer and cooler so that it is convenient to subdivide the stage according to the prevailing climate of the time. In the first, Preboreal, phase the frost tundra of the country south of the ice began to be covered with growths of the dwarf or arctic birch, a shrub with stems and branches generally spreading over the ground and making a bush only a couple of feet or so high in sheltered places; it is a characteristic plant of the arctic and high mountains. It was followed by the tree birches spreading to make a forest so that in the following Boreal phase, when they were joined by pine and hazel, the forest cover was complete. During the Boreal phase the melting of the ice brought a rapid rise in the level of the sea which finally cut through the Strait of Dover about 7,000 years ago, whereas the southern part of the North Sea and eastern end of the English Channel had been dry at the beginning of the Pre-boreal. At the same time the climate became several degrees warmer than at present, producing the Atlantic phase, during which the forest cover was enriched by the addition of oak, and alder. Thereafter the climate became cooler about 4,500 years ago, introducing the Sub-boreal phase, and the forest was further enriched by ash, elm and lime. A minor rise in sea level marked the end of the Sub-boreal phase about 2,250 years ago, and the climate entered the Sub-atlantic phase that we endure at the present day.

      At about the end of the Devensian glaciation the upper palaeolithic culture that man had evolved through many grades during the middle Pleistocene, was succeeded by the mesolithic culture characterised by the small flint artifacts called ‘microliths’. Of the several known mesolithic occupational sites, that at Starr Carr near Scarborough in Yorkshire has been meticulously excavated by Professor J. Clark and his colleagues, who have been able to draw a picture of the life of the inhabitants, and of the fauna and flora.³⁵ The site was occupied about 9,500 years ago as a winter hunting camp by three or four families of nomadic people; the settlement lay at the edge of a lake in the Vale of Pickering, with closed birch forest on the hill rising behind and willows along the reedy shore. The people lived on a platform of birch brushwood and were occupied not only in hunting and gathering roots of reeds and bog-bean, but also in knapping flint to make tools, some of which were used for making barbed spearheads from slivers cut from the antlers of red deer. The bones and antlers they left on the site show that they lived mostly on the flesh of red deer, but that they also killed roe deer, elk, aurochs and wild boar in lesser numbers. They had no domestic animals, and did not cultivate any crops. The remains of other mammals show that the fauna included the pine marten, fox, wolf, badger, hedgehog, hare and beaver.

      Fig. 5. The Flandrian succession in the British Isles after the ice of the Devensian glaciation melted.

      This late Pre-boreal fauna shows that forest animals, the deer and aurochs, had replaced the tundra-living mammals such as the reindeer, bison and wild horse while the birch forest increased with the rise in temperature. Until the rise in sea level at the end of the Preboreal phase, although Ireland had long been separated from Great Britain, the site of the southern part of the North Sea was dry land with the coast line extending from Flamborough Head to Jutland with a northern loop including the Dogger Bank. At the same time the English Channel extended no further east than Beachy Head.

      The mesolithic people remained in occupation for over five thousand years, during which the sea level rose and cut off the British Isles from the continent. About 4,500 years ago the neolithic people arrived, migrating across the sea from the east, and soon completely replaced the mesolithic culture with their own. Throughout the many hundreds of thousands of years of the preceding part of the Pleistocene the palaeolithic and later the mesolithic people were no more than part of the fauna, and produced no appreciable alteration in the environment. They were plant gatherers and hunters and, though they may have contributed towards the extinction of some of the large mammals such as the mammoth, their influence on the composition of the fauna was in general negligible. They probably had to work hard to earn their living in the British Isles, but further south on the continent, where the climate was milder and food abundant, they appear to have satisfied their wants more easily so that they had enough leisure to make paintings on the walls of caves, to make sculptures and carvings, and to engrave stones and bone artifacts with abstract and representational designs.