LIFE OF THE PAST
James O. Farlow, editor
HADROSAURS
Edited by DAVID A. EBERTH and DAVID C. EVANS
Editorial Assistant PATRICIA E. RALRICK
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Library of Congress Cataloging-in-Publication Data
Hadrosaurs / edited by David A. Eberth and David C. Evans.
pages cm. – (Life of the past)
Includes bibliographical references and index.
ISBN 978-0-253-01385-9 (cloth : alk. paper) – ISBN 978-0-253-01390-3 (ebook)
1. Hadrosauridae. 2. Hadrosauridae – Anatomy.
3. Hadrosauridae – Geographical distribution.
4. Dinosaurs. I. Eberth, David A. II. Evans, David C. (David Christopher), [date]
QE862.O65H33 2014
567.914 – dc23
2014011885
1 2 3 4 5 20 19 18 17 16 15
To David Weishampel and all those, from J. Leidy onward, who have contributed to our knowledge of hadrosaurs.
In particular, we recognize the efforts of Derek J. Main, a tireless promoter of Earth Science education and research. We value his contribution to this volume and mourn his all-too-soon passing.
Those animals of other days will give joy and pleasure to generations yet unborn.
Charles H. Sternberg
C
Contributors
Andrey Atuchin, Palaeontological Laboratory of the Institute of Geology and Nature Management, Far East Branch, Russian Academy of Sciences, per. Relochny 1, 675000 Blagoveschensk, Russia
Karl T. Bates, Department of Musculoskeletal Biology II, Institute of Aging and Chronic Disease, University of Liverpool, Sherrington Buildings, Ashton Street, Liverpool, U.K. L69 3GE
Paul M. Barrett, Department of Earth Sciences, Natural History Museum, Cromwell Road, London, U.K. SW7 5BD
Phil R. Bell, School of Environmental and Rural Science, University of New England, Armidale 2351, NSW, Australia
Uwe Bergmann, Stanford Synchrotron Radiation Lightsource, Stanford University, Menlo Park, California 94305
Ivan Y. Bolotsky, Research Center for Paleontology and Stratigraphy, Jilin University, Changchun 130061, China
Yuri L. Bolotsky, Palaeontological Laboratory of the Institute of Geology and Nature Management, Far East Branch, Russian Academy of Sciences, per. Relochny 1, 675000 Blagoveschensk, Russia
Kirstin S. Brink, Department of Ecology and Evolutionary Biology, University of Toronto Mississauga, 3359 Mississauga Road N., Mississauga, Ontario L5L 1C6
Donald B. Brinkman, Royal Tyrrell Museum of Palaeontology, Box 7500, Drumheller, Alberta T0J 0Y0
Caleb M. Brown, Royal Tyrell Museum of Paleontology, Box 7500, Drumheller, Alberta T0J 0Y0
Michael Buckley, Faculty of Life Sciences, University of Manchester, Manchester, U.K. M13 9PL
Michael E. Burns, Department of Biological Sciences, University of Alberta, Edmonton, Alberta T6G 2E9
Nicolás E. Campione, Departments of Earth Science and Organismal Biology, Uppsala University, Box 256, 751 05 Uppsala, Sweden
Rodolfo A. Coria, CONICET–University of Rio Negro–Museo Carmen Funes, Av. Córdoba 55 (8318) Plaza Huincul, Neuquén, Argentina
Philip J. Currie, Department of Biological Sciences, Zoo. 413, Biological Sciences Building, University of Alberta, Edmonton, Alberta T6G 2E9
Fabio M. Dalla Vecchia, Grup de Recerca del Mesozoic, Institut Catalá de Paleontologia Miquel Crusafont (ICP), Escola Industrial 23, E-08201 Sabadell, Spain
Peter Dodson, School of Veterinary Medicine, University of Pennsylvania, 3800 Spruce Street, Philadelphia, Pennsylvania 19104-6045
David A. Eberth, Royal Tyrrell Museum of Palaeontology, Box 7500, Drumheller, Alberta T0J 0Y0
Victoria M. Egerton, School of Earth, Atmospheric and Environmental Sciences, University of Manchester, Manchester, U.K. M13 9PL
Gregory M. Erickson, Department of Biological Science, Florida State University, Tallahassee, Florida, 32306-4295
David C. Evans, Department of Natural History, Royal Ontario Museum, 100 Queen’s Park, Toronto, Ontario M5S 2C6
Federico Fanti, Dipartimento di Scienze della Terra e Geologico-Ambientali, Universitá di Bologna, Italy
Andrew A. Farke, Raymond M. Alf Museum of Paleontology, 1175 West Baseline Road, Claremont, California 91711
Rodrigo Gaete, Museu de la Conca Dellá, Carrer del Museu 4, Isona, E-25650, Spain
Àngel Galobart, Grup de Recerca del Mesozoic, Institut Catalá de Paleontologia Miquel Crusafont (ICP), Escola Industrial 23, E-08201 Sabadell, Spain
Albert Garcia Sellés, Grup de Recerca del Mesozoic, Institut Catalá de Paleontologia Miquel Crusafont (ICP), Escola Industrial 23, E-08201 Sabadell, Spain
Terry A. Gates, Department of Biological Sciences, North Carolina State University, Raleigh, NC 27695; North Carolina Museum of Natural Sciences, Raleigh, NC 27601
Michael A. Getty, Denver Museum of Nature and Science, 2001 Colorado Boulevard, Denver, Colorado 80205-5732
Pascal Godefroit, Department of Palaeontology, Royal Belgian Institute of Natural Sciences,